1. Abeysingha DA, Dinesh D, Roopesh MS, Warkentin TD, Thilakarathna MS. (2024). The effect of cold plasma seed treatments on nodulation and plant growth in pea (Pisum sativum) and lentil (Lens culinaris). Plasma Processes and Polymers. 2024: e2400015.
  2. Rathor P, Upadhyay P, Ullah A, Gorim LY, Thilakarathna MS. (2024). Humic acid improves wheat growth by modulating auxin and cytokinin biosynthesis pathways. AoB PLANTS, plae018.
  3. Rathor P, Rouleau V, Chen G, Gorim LY, Thilakarathna MS. (2024). Humalite enhances the growth, grain yield, and protein content of wheat by improving soil nitrogen availability and nutrient uptake. Journal of Plant Nutrition and Soil Science, 1-13.
  4. Abeysingha DA, Dhaliwal HK, Du L, De Silva C, Szczyglowski K, Roopesh MS, Thilakarathna MS. (2024). The potential of cold plasma-based seed treatments on legume-rhizobia symbiotic nitrogen fixation: A review. Crops. 2024. 4: 95-114.


  1. De Silva C, Rathor P, Thilakarathna MS. (2023).Effects of drought stress on red clover-grass mixed stands compared to grass monoculture stands in nitrogen-deficient systems. Nitrogen. 4, 382-396
  2. Dollete D, Lumactud RA, Carlyle C, Szczyglowski K, Hill B. and Thilakarathna MS. (2023). Effect of drought stress on symbiotic nitrogen fixation, soil nitrogen availability and soil microbial diversity in forage legumes. Plant and Soil
  3. Rathor P, Gorim LY, Thilakarathna MS. (2023). Plant physiological and molecular responses triggered by humic based biostimulants-A way forward to sustainable agriculture. Plant and Soil, 492: 31-60.
  4. Liyanage DK, Torkamaneh D, Belzile F, Balasubramanian P, Hill B, Thilakarathna MS. (2023). The genotypic variability among short-season soybean cultivars for nitrogen fixation under drought stress. Plants 12: 1004.
  5. Lumactud RA, Dollete D, Liyanage DK, Szczyglowski K, Hill B. and Thilakarathna MS. (2023). The effect of drought stress on nodulation, plant growth, and nitrogen fixation in soybean during early plant growth. Journal of Agronomy and Crop Science. 209: 345-354  
  6. Enesi RO, Dyck M, Chang S, Thilakarathna MS, Fan X, Strelkov S, Gorim L.(2023).  Liming remediates soil acidity and improves crop yield and profitability-A Metaanalysis. Frontiers in Agronomy, 5, 1194896.


  1. Lumactud R, Gorim LU, Thilakarathna MS. (2022). Impacts of humic-based products on the microbial community structure and functions toward sustainable agriculture. Frontiers in Sustainable Food Systems. 6: 977121.
  2. Liyanage, D.K., Chathuranga, I., Mori, B. and Thilakarathna M.S. (2022). A simple, semi-automated, gravimetric method to simulate drought stress on plants. Agronomy: 12: 349.
  3. Ampong, K., Thilakarathna, M.S. and Gorim, L.Y. (2022). Understanding the role of Humic acids on crop performance and soil health. Frontiers in Agronomy. 4: 848621.


  1. Thilakarathna M.S., Torkamaneh D., Bruce R.W., Rajcan I., Chu G., Grainger C.M., Szczyglowski K., Hill, B. and Raizada, M.N. (2021). Testing whether pre-pod-fill symbiotic nitrogen fixation in soybean is subject to drift or selection over 100 years of soybean breeding. Frontiers in Agronomy: 3: 725813.
  2. Thilakarathna M.S. and Cope K. (2021). Split-root assays for studying legume-rhizobia symbioses, rhizodeposition, and belowground nitrogen transfer in legumes. Journal of Experimental Botany.
  3. Iqdiam B.M., Feizollahi E., Arif M.F., Jeganathan B., Vasanthan T., Thilakarathna M.S. and Roopesh M.S. (2021). T-2 and HT-2 toxin degradation by atmospheric cold plasma and its impact on quality changes and germination of wheat grains. Journal of Food Science. 86: 1354−1371.


  1. Feizollahi E., Iqdiam B., Vasanthan T., Thilakarathna M.S. and Roopesh, M.S. (2020). Effects of atmospheric-pressure cold plasma treatment on deoxynivalenol degradation, quality parameters, and germination of barley grains. Applied Sciences. 10: 3530.


  1. Thilakarathna M.S. and Raizada M.N. (2019). A biosensor-based assay shows defoliation triggers rapid release of glutamine from nodules and young roots of forage legumes. Phytobiomes Journal. 3: 85-91.
  2. Thilakarathna M.S., Chapagain T., Ghimire B., Pudasaini R., Tamang B., Gurung K., Choi K., Rai L., Magar S., BK B., Gaire S. and Raizada M.N. (2019). Evaluating the effectiveness of rhizobia inoculants and micronutrients as technologies for Nepalese common bean smallholder farmers in the real world context of highly variable terrace environments and indigenous farming practices. Agriculture. 9(1): 20.


  1. Thilakarathna M.S. and Raizada M.N. (2018). Challenges in using precision agriculture to optimize symbiotic nitrogen fixation in legumes: progress, limitations, and future improvements needed in diagnostic testing. Agronomy. 7: 78.
  2. Thilakarathna M.S. and Raizada M.N. (2018). Visualizing glutamine accumulation in root systems involved in the legume-rhizobia symbiosis by placement on agar embedded with companion biosensor cells. Phytobiomes Journal. 2: 117-128.
  3. Thilakarathna M.S., Papadopoulos Y.A., Grimmett M., Fillmore S.A., Crouse M. and Prithiviraj B. (2018). Red clover cultivars show nitrogen fixing advantage during the early stages of seedling development. Canadian Journal of Plant Science. 98(3): 517-526.


  1. Thilakarathna M.S., Moroz N. and Raizada M.N. (2017). A biosensor-based leaf punch assay for glutamine correlates to symbiotic nitrogen fixation measurements in legumes to permit rapid screening of rhizobia inoculants under controlled conditions. Frontiers in Plant Science. 8: 1714.
  2. Thilakarathna M.S., Janovicek K., Johnson P., Falk D., Navabi A. and Deen B. (2017). Frost seeding increases spring cereal yield. Canadian Journal of Plant Science. 97(3): 486-493.
  3. Thilakarathna M.S. and Raizada M.N. (2017). A meta-analysis of the effectiveness of diverse rhizobia inoculants on soybean traits under field conditions. Soil Biology and Biochemistry. 105: 177-196.